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I think, usually, you just cut a small piece of the bottom tier.
That's what I was thinking, but you'd be amazed how scarce information is on google about this!
we cut the corner of the bottom tier. but make sure you have a plate and a fork handy. we didn't have a fork nearby so i fed mr. sew with the big fat knife. :P they told me later i was supposed to use my fingers. bah!
We cut a teeny little 1" by 1" piece from the back of the cake and used our fingers to feed each other. There was no way we were going to be able to take an actual slice out of the cake because each layer took up almost all of the layer below it. Ahhh well. It eliminated my cake to the face fears because we only had a tiny mouthful each.
ok so first: who is cutting your cake? Normally the caterer takes care of cutting the slices for the guests. They'll know how to do this - if not you can find a guide here
http://www.simplycakesnc.com/Gallery.htm (click how to cut tiered/wedding cake pdf)
As far as the bride/groom photo opp cutting the cake - normally its best to cut a small piece from the bottom tier. It doesn't matter how big or small it is. Just maybe a few inches in. The staff will just work around whatever slice you've made later. Make sure you've got the cutting set, a plate, forks if wanted and a napkin handy!
And interesting fact, bridal superstition says that he should feed you first! I forget why this is though.
@Rgeddy: The caterer is definitely cutting up the cake for our guests!
@sewing: That's hilarious! I'll have to remember the fork and plate - it should also cut down on any desire FH has to smear it all over my face. I already told him that would be grounds for immediate annullment 
To add to the details...I read that his hand goes on top. Does that even matter? Also, I read that he feeds you first, as someone else mentioned.
You know, wasn't there a Bee who was very adamant about proper holding of the knife during cake cutting? Can't for the life of me remember who it was, but she was very serious about the wife's hand being on the knife and the husband's hand being over the wife's. Can't remember who she was but darned if I don't remember the proper way to cut a cake now. ;)
The woman's hand is on bottom because while she "rules the house" the husband "protects" it. And he feeds you first as a sign of providing for you.
Just FYI :D
We cut the from the bottom tier. We didn't really think about whose hand was on top though.
We just cut a normal looking cake slice (though thinner) in the bottom tier. The caterer cut around it!
We cut all way across the top tier, effectively cutting 2 slices, the back one fell off and slipped down the cake. It was really funny!
You can do whatever you want. There are no rules. Most people cut a chunk from the bottom tier, even though most bakers and magazines say to do the middle but it really doesn't matter.
I don't care where we cut it and how we eat it, I just want to eat it. A slice. A whole one. Maybe more than one.
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How do you do it? I mean, the bottom layer is going to be used for serving our guests, so I want to make sure I don't mangle the cake!
Do we just cut a little sliver of the tier that is visible?
Who would have thought that at 2.5 weeks to go, this would be causing me the most amount of stress?!?