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Ya, usually filling is in between layers of cake. Like cake cream, whipping cream, jams etc. :)
@toothfairyb: Ahh. So instead of icing? Like when I make a birthday cake I ice the entire first layer and then put the second on top so when you cut it there is icing/frosting inbetween layers.. but in this case there is only icing on the outside of the cake right?
Filling is in between the layers. Go to a local grocery bakery and get a cake of your choice and it will have some type of filling in the middle. As an example, if you get a carrot cake, it will have cream cheese frosting in the middle as well on the outside. The filling can be anything you want but if you are having a two layer cake (which is different from a tier) as nearly any baker will prepare unless specifically told otherwise via custom order, then it will have filling to make two (or more) layers stick together and in that case is not optional.
Thanks y'all. I think I'm going to look for a baker that offers this! Sounds yummy.
If nothing else, check grocery store bakeries since most offer yummy cakes. If they don't offer beyond vanilla for their wedding cakes, you can get a variety of regular round cakes (that are not classified as wedding cakes) from them and serve a cake buffet, which is cheaper in the end, even though grocery store wedding cakes are inexpensive to begin with.
I'd definitely check with the baker! When I get told a cake has filling I assume that it looks a bit like a Victoria Sponge cake. So two cakes sandwiching jam and/or cream and then on top of that a smaller cake of the same and then the whole thing covered on the outside in icing so you can't see the jam and cream inbetween :)
@Selene221: Well I have a specific cake look in mind so I really want to do that... even though it won't be too big as our wedding guest list is pretty tiny (49 right now and a bunch will probably not come since it's in another state)..
I am having trouble because we live in TX and are marrying in Iowa and we won't be able to go to Iowa until the week before the wedding (arriving late Saturday, marrying the following Saturday)... and I know that HERE anyway, the grocery store cakes are pretty consistently nasty, dry, gross. :( And that's several different grocery stores plus wal-mart's bakery. All yuck. Might as well just frost some stacked up sponges! 
@littlemissmoo: I will. I was just surprised when on his website he didn't mention fillings at all. Just cake flavors (and not very many!) and some pictures of his cakes. He has tons of high ratings on various online local ratings pages for his cakes being awesome.. but his flavors were so limited and fillings not mentioned at all.
I just really want to deal with someone (ideally) where I can SEE pics of their work online and their options and prices and reviews but I have to know that they'll deliver to our wedding location too which is a good hour from practically ANY large city in any direction. Headache!
Call or email and find out what his full menu of options are and if he is willing to anything custom for you. Many bakers are willing to do custom orders but they won't know what you want until you ask, even if it is something that is not on their menu.
@MsInterpret:My initial thought to the fact the baker doesn't have filling choices means to me that he doesn't offer them. Not all bakers will. But you could get, for example, a chocolate cake with coffee icing or something - it's essentially the same thing.
Filling is usually in between the layers. A wedding cake tier, it actually 2 layers of "normal" cake. So the filling will be between the two layers, of the tiers! We have having a strawberry filling on our wedding cake, but the frosting is just normal buttercream.
Hope that helps. I have a picture that shows the filling, I will try and find it.
Check with your venue for bakers--they may have some recommendations or even have a pastry chef on staff.
2 layers + filling=1 tier
Oooh, our cake will have three layers per tier, and filling between each layer... did I score? 
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I don't understand what you guys mean when you refer to a cake with "filling". Do you mean in between the tiers/layers, or somewhere inside the cake layer itself?
I know, weird question but I haven't been to a wedding in ages and the last one was just a cake, with icing in between the tiers... and I've only spoken to one baker so far (have a hard time finding bakers in the area we're getting married who have a web presence!) and he didn't offer fillings of any kind.